Stenopogoninae sensu Dikow 2009a

Scleropogon duncani , image © Torsten Dikow.


  • Autapomorphies: None.
  • Additional apomorphic character states:
    – facial swelling and mystax extending over lower facial half (4, 7)
    – postmentum ventrally with median groove in distal part (19)
    – all ommatidia same size (46)
    – postpedicel cylindrical throughout (54)
    – setae on anteroventral prothoracic tibiae absent (105)
    – female spermathecal reservoir formed by more or less expanded and coiled ducts (180)
  • Notes:
    Stenopogoninae sensu previous authors has been suspected to be non-monophyletic and used to be the largest taxon of Asilidae containing 118 genera (Geller-Grimm 2004) of which 37 are included in Dikow (2009). It was divided into 10 subordinate taxa to accommodate this diversity in a meaningful way by previous authors. In the new delimitation of Stenopogoninae, only three of the subordinate taxa are recognised by Dikow (2009) and the remaining taxa are either placed as a higher-level taxon, i.e., Bathypogoninae, Phellinae, Tillobromatinae, and Willistonininae, or transferred to another higher-level taxon, i.e., Acronychini to Leptogastrinae, Ceraturgini and Cyrtopogonini to Brachyrhopalinae.
  • Enigmomorphini: Connomyia, Creolestes, Daspletis, Microstylum, Prolepsis

  • Autapomorphies: None.
  • Additional apomorphic character states:
    – maxillary stipites fused medially, but V-shaped indentation present (14)
    – maxillae proximally high and distal tip distinctly narrower (28)
    – regular setae form discal scutellar setae (99)
    – female spermathecae occupying at most three posterior segments (182)
  • Notes:
    The Enigmomorphini was originally described by Hull (1962) to accommodate the Neotropical genus Enigmomorphus, and later expanded by Artigas & Papavero (1991) to include 14 genera. The New World genus Archilestris is transferred to the Dasypogoninae and the two Afrotropical genera Connomyia and Daspletis are transferred here to Enigmomorphini.
  • Plesiommatini: Plesiomma

  • Autapomorphies: None.
  • Additional apomorphic character states:
    – vertex between compound eyes sharply depressed (3)
    – facial swelling extending over entire face (4)
    – mystax restricted to lower facial margin (7)
    – dorsal flange medially on prementum absent (22)
    – hypopharynx seta-like spicules spaced far apart (33)
    – pharyngeal pump with entire sclerotised ring on lateral margin and circular in shape (40, 41)
    – anterior tentorial pits well developed and conspicuous anteromedially located (44)
    – frons at antennal insertion markedly approximating medially at level of antennal insertion (48)
    – regular setae on lateral margin of frons dorsal to antennal insertion (no macrosetae) (49)
    – regular setae forming postpronotum setae (no macrosetae) (70)
    – scutellum small and mesopostnotum visible in dorsal view (97)
    – apical scutellar setae absent (98)
    – regular setae forming setation on ventral metathoracic femora (116)
    – T2 longer than wide, but less than twice as long (153)
    – female common spermathecal duct long, extending beyond tip of furca (177)
    – male lateral ejaculatory process present and formed by a large cylindrical sclerite (213)
  • Notes:
    The Plesiommatini was described by Artigas & Papavero (1991) to accommodate the three Neotropical genera Cystoprosopa, Dapsilochaetus, and Plesiomma.
  • Stenopogonini: Gonioscelis, Ospriocerus, Scleropogon, Stenopogon

  • Autapomorphies: None.
  • Additional apomorphic character states:
    – head circular, as wide as high, in anterior view (1)
    – maxillary stipites divided medially (14)
    – ventral margin of postmentum entirely smooth (19)
    – dorsoposterior margin of cibarium with only one ridge connecting cornua and cornua not originating from ridge (35)
    – cornua developed in anteroposterior orientation (37)
    – prosternum and proepisternum separated and prosternum triangular with pointed dorsal tip (74)
    – cell m3 closed (135)
    – R2+3 apex arching sharply anteriorly in 90 degrees angle (142)
    – male hypopygium not rotated (195)
  • Notes:
    The Stenopogonini has been restricted by Dikow (2009) to only those robber flies with a circular head and with the character combination mentioned above.
  • Genera not placed within above tribal taxa:

  • Ancylorhynchus, Scylaticus